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Soil is unconsolidated rock particles in combination with organic matter.
Soil is vital to all life on Earth because it supports the growth of plants, which supply food and oxygen and absorbs carbon dioxide and nitrogen.
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Soils vary widely in composition and structure from place to place. Soils are formed through the weathering of rock and the breakdown of organic matter. Weathering is the action of wind, rain, ice, sunlight and biological processes on rocks, which breaks them down into small particles. The proportions of minerals and organic matter determine the structure and other characteristics of a particular soil.
Soils can be divided into two general layers or strata: topsoil, the topmost layer, where most plant roots, microorganisms, and other animal life are located, and subsoil, which is deeper and often more dense and less rich in organic matter.
Water and air are also components of most soils. Air, trapped in spaces between soil particles, and water, trapped in spaces and on the surface of particles, comprises about half of the soil by volume. Both are important to plant growth and other life in the soil profile of a particular ecosystem.
The rock and mineral content of soil is categorized according to particle size as sand (coarsest), silt or clay (finest); the ratio of these particles to a great degree determines the soil classification and characteristics.
Soil serves as a habitat soil organisms varying in size from microorganisms to small animals. The character of soil is intricately tied to bioturbation and the biochemical functions performed by soil organisms.
Former soils which become buried below the effects of organisms are called paleosols.
Soil develops naturally over time through the action of plants, animals, and weathering. Soil is also affected by human habitation. People can alter soil to make it more suitable for plant growth through the addition of organic material and natural or synthetic fertilizer, and by improving its drainage or water-retaining capacity. Human actions also can degrade soil through the depletion of nutrients, pollution, contamination, and compaction, and by increasing the rate of erosion, which is the relocation of soil through the movement of water or wind.
An example of soil development from bare rock occurs on recent lava flows in warm regions under heavy and very frequent rainfall. In such climates plants become established very quickly on basaltic lava, even though there is very little organic material. The plants are supported by the porous rock becoming filled with nutrient bearing water, for example carrying dissolved bird droppings or guano. The developing plant roots themselves gradually break up the porous lava and organic matter soon accumulates but, even before it does, the predominantly porous broken lava in which the plant roots grow can be considered soil.
Weathering releases ions such as Potassium (K+) and Magnesium (Mg2+) into the soil solution. Some of these elements (as ions) are taken up by bacteria, fungi and plants. The remaining portion can form secondary minerals, be chelated into organic complexes or be adsorbed into ion exchange complexes. Anion exchange complexes affect negatively charged ions (phosphate) and compounds. Anion exchange surfaces occur most typically in humus. Cation exchange complexes affect positively charged ions. Cation exchange surfaces are typically clay minerals such as montmorillonite and organic materials such as humus. When the level of ions is relatively low in the soil solution, equilibrium processes convey ions into solution, where they satisfy demand for nutrients by plants, bacteria and fungi.
The pH level in soil affects the activity and availability of ionic nutrients (examples are Ca2+, Mg2+, K+, Na+) and non-nutrients (H+, Al3+). Nutrient uptake is highest in a neutral pH range of 5.5 to 8.2. At pH levels below 5.0, increased aluminum activity can have a toxic affect, exacerbating reduced nutrient availability. Additionally, Ca2+, Mg2+, K+, Na+ can be displaced by H+ and Al3+. Subsequent leaching can result in lower soil fertility and productivity. At elevated soil pH levels nutrient availability is limited, especially for zinc and phosphorus. Additionally, differential removal of cations can result in elevated Na+ relative to Ca2+ and Mg2+ with a deleterious affect on soil structure, permeability and tilth. Contributors to soil acidification include "acidic" parent material (granite), plant root exudates, decomposition of certain types of organic residue (pine needles), chemical changes that occur when perennially wet sediments are dried, acidifying fertilizers (anhydrous ammonia, ammonium sulfate), and natural rain as well as acid rain phenomena. Sources of alkalinity include "basic" parent material (serpantine, limestone) and airborne soil particulates from alkaline areas. To raise soil pH, farmers can apply alkaline materials such as lime. To lower soil pH, farmers can apply acid-forming materials such as elemental sulfur. To increase calcium content in an alkaline soil, farmers can apply gypsum.
Although the elements nitrogen, potassium and phosphorus, which are necessary for plant growth, may be abundant in soil, only a fraction of these elements may be in a chemical form which plants can use.
Processes such as the nitrogen cycle and carbon cycle continually exchange nitrogen and carbon nutrients between the soil and atmosphere. The raw products are initially present as gases in the atmosphere. In nitrogen fixation, atmospheric nitrogen is converted to plant available forms. In nitrogen mineralization, proteins and other organic forms are converted into mineral, plant available forms: NH4+ and NO3-. In nitrification, NH4+ is converted into the more usable NO3-. While NH4+ is especially important to young plants and early in the growing season, NO3- is the dominant form of nitrogen taken up by plants. NO3- moves to plants by mass transport and needs transpiration to drive uptake.
The organic component of soils originate in plant debris (such as fallen leaves), animal excreta, and other decomposing organic materials. These materials, when broken down, form humus, a dark, nutrient-rich material. Chemically, humus is composed of very large molecules including esters of carboxylic acid, phenolic compounds, and derivatives of benzene. Organic material in soil provides nutrients necessary for plant growth. Organic material also contributes to water retention, drainage ability, and oxygenation of soil.
If oxygen enters a wet soil, because of lowered ground water table, organic matter in the soil will be broken down further by oxidation, which can lead to subsidence. An example of this can be seen in soils in the Everglades region of Florida, which have been drained by canals for agriculture, primarily sugar production. Originally very high in organic content, oxygenation and compaction have led to breakdown of the soil structure and nutrient content, and degradation of the soil's ability to support continued high crop yields.
The diffusion of dissolved oxygen in saturated soils is slower than in unsaturated soils. Wetland (also referred to as hydric) soils form due to soil microbial cellular respiration in excess of soil oxygen supply, resulting in oxygen depletion. Anaerobic soil chemistry results, which creates a reducing environment. This eliminates plants and creatures not adapted for life in saturated soil conditions.
Biological soil crusts are formed by living organisms and their by-products, creating a surface crust of soil particles bound together by organic materials.
Nutrition is the study of the relationship between diet and states of health and disease. It is defined as the study of food. Absence of adequate nutrients can cause certain diseases to take hold that can potentially result in death.
Between the extremes of optimal health and death from starvation or malnutrition, there is an array of disease states that can be caused or alleviated by changes in diet. Deficiencies, excesses and imbalances in the diet can produce negative impacts on health, which may result in diseases such as scurvy, obesity or osteoporosis. Also, excess ingestion of elements that have no apparent role in health (e.g. lead, mercury, PCBs, dioxins) may have toxic and potentially lethal effects depending on dose.
The science of nutrition attempts to understand how and why specific aspects of diet have specific influences on health.
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The human body comprises chemical compounds such as water, amino acids (proteins), fatty acids (lipids), nucleic acids (DNA/RNA), and carbohydrates (e.g. sugars). These compounds in turn consist of elements such as carbon, hydrogen, oxygen, nitrogen, and phosphorus, and may or may not contain minerals such as calcium, iron, and zinc. Minerals also ubiquitously occur in the form of salts and electrolytes. All of these chemical compounds and elements occur in various forms and combinations (e.g. hormones/vitamins, phospholipids, hydroxyapatite), both in the human body and in organisms (e.g. plants, animals) that humans eat.
The human body must necessarily comprise those elements that humans eat and absorb into the bloodstream. The digestive system, except in the unborn fetus, is the first step in helping to make the different chemical compounds and elements in food available for the trillions of cells of the body. In the digestive process of an average adult, about seven (7) litres of liquid, known as digestive juices, exit the internal body and enter the lumen of the digestive tract. The digestive juices help break chemical bonds between ingested compounds as well as modulate the conformation and/or energetic state of the compounds/elements. Yet many compounds/elements are absorbed into the bloodstream unchanged, though the digestive process helps to release them from the matrix of the foods where they occur. Any unabsorbed matter is eliminated in the feces. Only a minimal amount of digestive juice is eliminated this way; the intestines reabsorb most of it otherwise the body would rapidly dehydrate (hence the devastating effects of persistent diarrhea).
Study in this field must take into careful account the state of the body before ingestion and after digestion as well as the chemical content of both the food and the waste. The specific types of compounds and elements that are absorbed by the body can be determined by comparing the waste to the food. The effect that the absorbed matter has on the body can be determined by finding the difference between the pre-ingestion state and the post-digestion state. The effect may only be discernible after an extended period of time in which all food and ingestion must be exactly regulated and all waste must be analyzed. The number of variables (e.g. 'confounding factors') involved in this type of experimentation is very high. This makes scientifically valid nutritional study very time-consuming and expensive, which accounts for why a proper science of human nutrition is rather new.
In general, eating a variety of fresh, whole (unprocessed) foods has proven hormonally and metabolically favourable compared to eating a monotonous diet based on processed foods. In particular, fresh, whole foods provide higher amounts and a more favourable balance of essential and vital nutrients per unit of energy, resulting in better management of cell growth, maintenance, and mitosis (cell division) as well as of appetite and energy balance. A generally more regular eating pattern (e.g. eating medium-sized meals every 3 to 4 hours) has also proven more hormonally and metabolically favourable than infrequent, haphazard food intake.
Ill health can be brought about by an imbalance of nutrients, producing either an excess or deficiency which in turn affects body functioning in a cumulative manner. Moreover, because most nutrients are, in some way or the other, involved in cell-to-cell signalling (e.g. as building block or part of a hormone or signalling 'cascades'), deficiency or excess of various nutrients affects hormonal function also indirectly. Thus, because they largely regulate the expression of genes, hormones represent a link between nutrition and how our genes are expressed, i.e. our phenotype. The strength and nature of this link are continually under investigation, but observations especially in recent years have demonstrated a pivotal role for nutrition in hormonal activity and function and therefore in health.
Mineral and/or vitamin (tocotrienol and tocopherol) deficiency or excess may yield symptoms of diminishing health such as goitre, scurvy, osteoporosis, weak immune system, disorders of cell metabolism, certain forms of cancer, symptoms of premature aging, and poor psychological health (including eating disorders). The list goes on and on; for reference, see Modern Nutrition in Health and Disease by Shils et al.
As of 2005, twelve vitamins and about the same number of minerals are recognized as 'essential nutrients', meaning that they must be consumed and absorbed - or, in the case of vitamin D, alternatively synthesized via UVB radiation - to prevent deficiency symptoms and death. Certain vitamin-like substances found in foods, such as carnitine, have also been found essential to survival and health, but these are not strictly 'essential' to eat because the body can produce them from other compounds. Moreover, thousands of different phytochemicals have recently been discovered in food (particularly in fresh vegetables), which have many discovered and yet to be discovered properties including antioxidant activity (see below). Other essential nutrients include essential amino acids, choline and the essential fatty acids.
In addition to sufficient intake, an appropriate balance of essential fatty acids - omega-3 and omega-6 fatty acids - has been discovered to be crucial for maintaining health. Both of these unique "omega" long-chain polyunsaturated fatty acids are substrates for a class of eicosanoids known as prostaglandins. The omega-3 eicosapentaenoic acid (EPA) (which can be made in the body from the omega-3 essential fatty acid alpha-linolenic acid (LNA), or taken in through marine food sources), serves as building block for series 3 prostaglandins (e.g. weakly-inflammation PGE3). The omega-6 dihomo-gamma-linolenic acid (DGLA) serves as building block for series 1 prostaglandins (e.g. anti-inflammatory PGE1), whereas arachidonic acid (AA) serves as building block for series 2 prostaglandins (e.g. pro-inflammatory PGE1). Both DGLA and AA are made from the omega-6 linoleic acid (LA) in the body, or can be taken in directly through food. An appropriately balanced intake of omega-3 and omega-6 partly determines the relative production of different prostaglandins, which partly explains the importance of omega-3/omega-6 balance for cardiovascular health. In industrialised societies, people generally consume large amounts of processed vegetable oils that have reduced amounts of essential fatty acids along with an excessive amount of omega-6 relative to omega-3.
The rate of conversions of omega-6 DGLA to AA largely determines the production of the respective prostaglandins PGE1 and PGE2. Omega-3 EPA prevents AA from being released from membranes, thereby skewing prostaglandin balance away from pro-inflammatory PGE2 made from AA toward anti-inflammatory PGE1 made from DGLA. Moreover, the conversion (desaturation) of DGLA to AA is controlled by the enzyme delta-5-desaturase, which in turn is controlled by hormones such as insulin (up-regulation) and glucagon (down-regulation). Because different types and amounts of food eaten/absorbed affect insulin, glucagon and other hormones to varying degrees, not only the amount of omega-3 versus omega-6 eaten but also the general composition of the diet therefore determine health implications in relation to essential fatty acids, inflammation (e.g. immune function) and mitosis (i.e. cell division).
Several lines of evidence indicate lifestyle-induced hyperinsulinemia and reduced insulin function (i.e. insulin resistance) as a decisive factor in many disease states. For example, hyperinsulinemia and insulin resistance are strongly linked to chronic inflammation, which in turn is strongly linked to a variety of adverse developments such as arterial microinjuries and clot formation (i.e. heart disease) and exaggerated cell division (i.e. cancer). Hyperinsulinemia and insulin resistance (the so-called metabolic syndrome) are characterized by a combination of abdominal obesity, elevated blood sugar, elevated blood pressure, elevated blood triglycerides, and reduced HDL cholesterol. The negative impact of hyperinsulinemia on prostaglandin PGE1/PGE2 balance may be significant.
The state of obesity clearly contributes to insulin resistance, which in turn can cause type 2 diabetes. Virtually all obese and most type 2 diabetic individuals have marked insulin resistance. Although the association between overfatness and insulin resistance is clear, the exact (likely multifarious) causes of insulin resistance remain less clear. Importantly, it has been demonstrated that appropriate exercise, more regular food intake and reducing glycemic load (see below) all can reverse insulin resistance in overfat individuals (and thereby lower blood sugar levels in those who have type 2 diabetes).
Overfatness can unfavourably alter hormonal and metabolic status via resistance to the hormone leptin, and a vicious cycle may occur in which insulin/leptin resistance and overfatness aggravate one another. The vicious cycle is putatively fuelled by continuously high insulin/leptin stimulation and fat storage, as a result of high intake of strongly insulin/leptin stimulating foods and energy. Both insulin and leptin normally function as satiety signals to the hypothalamus in the brain; however, insulin/leptin resistance may reduce this signal and therefore allow continued overfeeding despite large bodyfat stores. In addition, reduced leptin signalling to the brain may reduce leptin's normal effect to maintain an appropriately high metabolic rate.
There is debate about how and to what extent different dietary factors - e.g. intake of processed carbohydrates, total protein, fat, and carbohydrate intake, intake of saturated and trans fatty acids, and low intake of vitamins/minerals - contribute to the development of insulin- and leptin resistance. In any case, analogous to the way modern man-made pollution may potentially overwhelm the environment's ability to maintain 'homeostasis', the recent explosive introduction of high Glycemic Index- and processed foods into the human diet may potentially overwhelm the body's ability to maintain homeostasis and health (as evidenced by the metabolic syndrome epidemic).
Antioxidants are another recent discovery. As cellular metabolism/energy production requires oxygen, potentially damaging (e.g. mutation causing) compounds known as radical oxygen species or free radicals may form. For normal cellular maintenance, growth, and division, these free radicals must be sufficiently neutralized by antioxidant compounds, such as certain vitamins (vitamin C, vitamin E, vitamin K and the aforementioned phytochemicals as well as other compounds, some of which the body itself produces. Different antioxidants are now known to function in a cooperative network, e.g. vitamin C can reactivate free radical-containing glutathione or vitamin E by accepting the free radical itself, and so on.
It is now also known that the human digestion system contains a population of a range of bacteria which are essential to digestion, and which are also affected by the food we eat. The role and significance of the intestinal bacterial flora is under investigation.
(Stub, please expand.) Nutrition is very important for improving sports performance. The most common means to improve performance through diet is the practice of eating large quantities of protein, usually red meat, when attempting to build muscle mass; its efficacy is doubtful, as daily protein intake even on a normal diet usually outweighs the amount of muscle protein which can be synthesized in a day.
To enhance their speed of muscle synthesis, athletes will focus a great deal on how to best accelerate their tissue recovery. Icing/heating the muscles to reduce swelling and increase blood flow, along with plenty of rest, and rehabilitative low-intensity exercising, stretching, and massage thereapy, along with plenty of sleep and nutrition (such as water and creatine, are instrumental in this.
Protein is a much less efficient source of the energy needed to build new muscle tissue than are fats and carbohydrates. Even so, athletes will usually overdo it, since they'd rather have leftover protein needed to be used for energy rather than not enough for ideal recovery and rebuilding of damage muscle tissue.
Lifespan may be somehow related to the amount of food energy consumed: this was first systematically investigated in the seminal study by Weidruch, et al. (1986). A pursuit of this principle of caloric restriction followed, involving research into longevity of those who reduced their food energy intake while attempting to optimize their micronutrient intake. Perhaps not surprisingly, some people found that cutting down on food reduced their quality of life so considerably as to negate any possible advantages of lengthening their lives. However, a small set of individuals persists in the lifestyle, going so far as to monitor blood lipid levels and glucose response every few months. See Calorie Restriction Society.
Underlying this research was the hypothesis that oxidative damage was the agent which accelerated aging, and that aging was retarded when the amount of carbohydrates (and thereby insulin release) was reduced through dietary restriction.
However, recent research has produced increased longevity in animals (and shows promise for increased human longevity) through the use of insulin uptake retardation. This was done through altering an animal’s metabolism to allow it to consume similar food-energy levels to other animals, but without building up fatty tissue. (Bluher et al, 2003)
This has set researchers off on a line of study which presumes that it is not low food energy consumption which increases longevity. Instead, longevity may depend on an efficient fat processing metabolism, and the consequent long term efficient functioning of our organs free from the encumbrance of accumulating fatty deposits. (Das et al, 2004) Thus, longevity may be related to maintained insulin sensitivity. However, several other factors including low body temperature seem to promote longevity also and it is unclear to what extent each of them contribute.
Antioxidants have recently come to the forefront of longevity studies which have included the FDA and Brunswick labs. In 2005 the FDA issued a statement recommending that Americans should be consuming 7,000 ORAC units daily or 12 full servings of fruit in order to curb the cancer epidemic. The dietary supplement industry has responded by shifting focus away from hormone replacements to “super” antioxidants such as Proleva which contain whole fruit extracts and ORAC scores near 5,000 units mark or two thirds of the new level set by the FDA.
Since the Industrial Revolution some two hundred years ago, the food processing industry has invented many technologies that both help keep foods fresh longer and alter the fresh state of food as they appear in nature. Cooling is the primary technology that can help maintain freshness, whereas many more technologies have been invented to allow foods to last longer without becoming spoiled. These latter technologies include pasteurisation, autoclavation, drying, salting, and separation of various components, and all appear to alter the original nutritional contents of food. Pasteurisation and autoclavation (heating techniques) have no doubt improved the safety of many common foods, preventing epidemics of bacterial infection. But some of the (new) food processing technologies undoubtedly have downfalls as well.
Modern separation techniques such as milling, centrifugation, and pressing have enabled upconcentration of particular components of food, yielding flour, oils, juices and so on, and even separate fatty acids, amino acids, vitamins, and minerals. Inevitably, such large scale upconcentration changes the nutritional content of food, saving certain nutrients while removing others. Heating techniques may also reduce food's content of many heat-labile nutrients such as certain vitamins and phytochemicals, and possibly other yet to be discovered substances. Because of reduced nutritional value, processed foods are often 'enriched' or 'fortified' with some of the most critical nutrients (usually certain vitamins) that were lost during processing. Nonetheless, processed foods tend to have an inferior nutritional profile than do whole, fresh foods, regarding content of both sugar and high GI starches, potassium/sodium, vitamins, fibre, and of intact, unoxidized (essential) fatty acids. In addition, processed foods often contain potentially harmful substances such as oxidized fats and trans fatty acids.
A dramatic example of the effect of food processing on a population's health is the history of epidemics of beri-beri in people subsisting on polished rice. Removing the outer layer of rice by polishing it removes with it the essential vitamin thiamin, causing beri-beri. Another example is the development of scurvy among infants in the late 1800's in the United States. It turned out that the vast majority of sufferers were being fed milk that had been heat-treated (as suggested by Pasteur) to control bacterial disease. Pasteurisation was effective against bacteria, but it destroyed the vitamin C.
As mentioned, lifestyle- and obesity-related diseases are becoming increasingly prevalent all around the world. There is little doubt that the increasingly widespread application of some modern food processing technologies has contributed to this development. The food processing industry is a major part of modern economy, and as such it is influential in political decisions (e.g. nutritional recommendations, agricultural subsidising). In any known profit-driven economy, health considerations are hardly a priority; effective production of cheap foods with a long shelf-life is more the trend. In general, whole, fresh foods have a relatively short shelf-life and are less profitable to produce and sell than are more processed foods. Thus the consumer is left with the choice between more expensive but nutritionally superior whole, fresh foods, and cheap, usually nutritionally inferior processed foods. Because processed foods are often cheaper, more convenient (in both purchasing, storage, and preparation), and more available, the consumption of nutritionally inferior foods has been increasing throughout the world along with many nutrition-related health complications.
Most Governments provide guidance on good nutrition, and some also impose mandatory labelling requirements upon processed food manufacturers to assist consumers in complying with such guidance. Current dietary guidelines in the United States are presented in the concept of a food pyramid. There is no apparent consisteny in science-based nutritional recommendations between countries, indicating the role of politics as well as cultural bias in research emphasis and interpretation.
Challenging issues in modern nutrition include:
'Artificial' interventions in food production and supply:
Sociological issues:
Research Issues:
For detailed information, see related entries in the following categories:
Food:
Health:
Research: